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Like all loyalty abilities, it can be activated only once per turn, during your main phase, when the stack is empty, and only if no other loyalty abilities of the planeswalker have been activated this turn

[4] Players deal damage to both creatures and to each other

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Then, thanks to Luminous Broodmoth, Uro, Titan of Nature's Wrath returns to the battlefield to draw us another card and gain us more life before dying again (since it wasn't escaped), which triggers Death's Oasis again to return another 2-CMC-or-less creature to our hand and mill a couple of more cards

It can be used at any time during combat, even after combat damage has been dealt